Geological Contemporaneity and Persistent Types of Life
Date: 2001-11-02T00:00:00+00:00
Keywords: Life (Biology); Fossils

The Project Gutenberg eBook of Geological Contemporaneity and Persistent Types of Life

This eBook is for the use of anyone anywhere in the United States and most other parts of the world at no cost and with almost no restrictions whatsoever. You may copy it, give it away or re-use it under the terms of the Project Gutenberg License included with this eBook or online at www.gutenberg.org. If you are not located in the United States, you will have to check the laws of the country where you are located before using this eBook.

Title: Geological Contemporaneity and Persistent Types of Life

Author: Thomas Henry Huxley

Release date: November 1, 2001 [eBook #2936]
Most recently updated: January 22, 2013

Language: English

Other information and formats: www.gutenberg.org/ebooks/2936

Credits: Produced by Amy E. Zelmer, and David Widger

*** START OF THE PROJECT GUTENBERG EBOOK GEOLOGICAL CONTEMPORANEITY AND PERSISTENT TYPES OF LIFE ***

GEOLOGICAL CONTEMPORANEITY AND PERSISTENT TYPES OF LIFE.

By Thomas H. Huxley

1

MERCHANTS occasionally go through a wholesome, though troublesome and not always satisfactory, process which they term "taking stock." After all the excitement of speculation, the pleasure of gain, and the pain of loss, the trader makes up his mind to face facts and to learn the exact quantity and quality of his solid and reliable possessions.

The man of science does well sometimes to imitate this procedure; and, forgetting for the time the importance of his own small winnings, to re-examine the common stock in trade, so that he may make sure how far the stock of bullion in the cellar—on the faith of whose existence so much paper has been circulating—is really the solid gold of truth.

The Anniversary Meeting of the Geological Society seems to be an occasion well suited for an undertaking of this kind—for an inquiry, in fact, into the nature and value of the present results of paleontological investigation; and the more so, as all those who have paid close attention to the late multitudinous discussions in which paleontology is implicated, must have felt the urgent necessity of some such scrutiny.

First in order, as the most definite and unquestionable of all the results of paleontology, must be mentioned the immense extension and impulse given to botany, zoology, and comparative anatomy, by the investigation of fossil remains. Indeed, the mass of biological facts has been so greatly increased, and the range of biological speculation has been so vastly widened, by the researches of the geologist and paleontologist, that it is to be feared there are naturalists in existence who look upon geology as Brindley regarded rivers. "Rivers," said the great engineer, "were made to feed canals"; and geology, some seem to think, was solely created to advance comparative anatomy.

Were such a thought justifiable, it could hardly expect to be received with favour by this assembly. But it is not justifiable. Your favourite science has her own great aims independent of all others; and if, notwithstanding her steady devotion to her own progress, she can scatter such rich alms among her sisters, it should be remembered that her charity is of the sort that does not impoverish, but "blesseth him that gives and him that takes."

Regard the matter as we will, however, the facts remain. Nearly 40,000 species of animals and plants have been added to the Systema Naturae by paleontologic research. This is a living population equivalent to that of a new continent in mere number; equivalent to that of a new hemisphere, if we take into account the small population of insects as yet found fossil, and the large proportion and peculiar organization of many of the Vertebrata.

But, beyond this, it is perhaps not too much to say that, except for the necessity of interpreting paleontologic facts, the laws of distribution would have received less careful study; while few comparative anatomists (and those not of the first order) would have been induced by mere love of detail, as such, to study the minutiae of osteology, were it not that in such minutiae lie the only keys to the most interesting riddles offered by the extinct animal world.

These assuredly are great and solid gains. Surely it is matter for no small congratulation that in half a century (for paleontology, though it dawned earlier, came into full day only with Cuvier) a subordinate branch of biology should have doubled the value and the interest of the whole group of sciences to which it belongs.

But this is not all. Allied with geology, paleontology has established two laws of inestimable importance: the first, that one and the same area of the earth's surface has been successively occupied by very different kinds of living beings; the second, that the order of succession established in one locality holds good, approximately, in all.

The first of these laws is universal and irreversible; the second is an induction from a vast number of observations, though it may possibly, and even probably, have to admit of exceptions. As a consequence of the second law, it follows that a peculiar relation frequently subsists between series of strata, containing organic remains, in different localities. The series resemble one another, not only in virtue of a general resemblance of the organic remains in the two, but also in virtue of a resemblance in the order and character of the serial succession in each. There is a resemblance of arrangement; so that the separate terms of each series, as well as the whole series, exhibit a correspondence.

Succession implies time; the lower members of a series of sedimentary rocks are certainly older than the upper; and when the notion of age was once introduced as the equivalent of succession, it was no wonder that correspondence in succession came to be looked upon as a correspondence in age, or "contemporaneity." And, indeed, so long as relative age only is spoken of, correspondence in succession 'is' correspondence in age; it is 'relative' contemporaneity.

But it would have been very much better for geology if so loose and ambiguous a word as "contemporaneous" had been excluded from her terminology, and if, in its stead, some term expressing similarity of serial relation, and excluding the notion of time altogether, had been employed to denote correspondence in position in two or more series of strata.

In anatomy, where such correspondence of position has constantly to be spoken of, it is denoted by the word "homology" and its derivatives; and for Geology (which after all is only the anatomy and physiology of the earth) it might be well to invent some single word, such as "homotaxis" (similarity of order), in order to express an essentially similar idea. This, however, has not been done, and most probably the inquiry will at once be made—To what end burden science with a new and strange term in place of one old, familiar, and part of our common language?

The reply to this question will become obvious as the inquiry into the results of paleontology is pushed further.

Those whose business it is to acquaint themselves specially with the works of paleontologists, in fact, will be fully aware that very few, if any, would rest satisfied with such a statement of the conclusions of their branch of biology as that which has just been given.

Our standard repertories of paleontology profess to teach us far higher things—to disclose the entire succession of living forms upon the surface of the globe; to tell us of a wholly different distribution of climatic conditions in ancient times; to reveal the character of the first of all living existences; and to trace out the law of progress from them to us.

It may not be unprofitable to bestow on these professions a somewhat more critical examination than they have hitherto received, in order to ascertain how far they rest on an irrefragable basis; or whether, after all, it might not be well for paleontologists to learn a little more carefully that scientific "ars artium," the art of saying "I don't know." And to this end let us define somewhat more exactly the extent of these pretensions of paleontology.

Every one is aware that Professor Bronn's 'Untersuchungen' and Professor Pictet's 'Traite de Paleontologie' are works of standard authority, familiarly consulted by every working paleontologist. It is desirable to speak of these excellent books, and of their distinguished authors, with the utmost respect, and in a tone as far as possible removed from carping criticism; indeed, if they are specially cited in this place, it is merely in justification of the assertion that the following propositions, which may be found implicitly, or explicitly, in the works in question, are regarded by the mass of paleontologists and geologists, not only on the Continent but in this country, as expressing some of the best-established results of paleontology. Thus:—

Animals and plants began their existence together, not long after the commencement of the deposition of the sedimentary rocks; and then succeeded one another, in such a manner, that totally distinct faunae and florae occupied the whole surface of the earth, one after the other, and during distinct epochs of time.

A geological formation is the sum of all the strata deposited over the whole surface of the earth during one of these epochs: a geological fauna or flora is the sum of all the species of animals or plants which occupied the whole surface of the globe, during one of these epochs.

The population of the earth's surface was at first very similar in all parts, and only from the middle of the Tertiary epoch onwards, began to show a distinct distribution in zones.

The constitution of the original population, as well as the numerical proportions of its members, indicates a warmer and, on the whole, somewhat tropical climate, which remained tolerably equable throughout the year. The subsequent distribution of living beings in zones is the result of a gradual lowering of the general temperature, which first began to be felt at the poles.

It is not now proposed to inquire whether these doctrines are true or false; but to direct your attention to a much simpler though very essential preliminary question—What is their logical basis? what are the fundamental assumptions upon which they all logically depend? and what is the evidence on which those fundamental propositions demand our assent?

These assumptions are two: the first, that the commencement of the geological record is coeval with the commencement of life on the globe; the second, that geological contemporaneity is the same thing as chronological synchrony. Without the first of these assumptions there would of course be no ground for any statement respecting the commencement of life; without the second, all the other statements cited, every one of which implies a knowledge of the state of different parts of the earth at one and the same time, will be no less devoid of demonstration.

The first assumption obviously rests entirely on negative evidence. This is, of course, the only evidence that ever can be available to prove the commencement of any series of phenomena; but, at the same time, it must be recollected that the value of negative evidence depends entirely on the amount of positive corroboration it receives. If A B wishes to prove an 'alibi', it is of no use for him to get a thousand witnesses simply to swear that they did not see him in such and such a place, unless the witnesses are prepared to prove that they must have seen him had he been there. But the evidence that animal life commenced with the Lingula-flags, 'e.g.', would seem to be exactly of this unsatisfactory uncorroborated sort. The Cambrian witnesses simply swear they "haven't seen anybody their way"; upon which the counsel for the other side immediately puts in ten or twelve thousand feet of Devonian sandstones to make oath they never saw a fish or a mollusk, though all the world knows there were plenty in their time.

But then it is urged that, though the Devonian rocks in one part of the world exhibit no fossils, in another they do, while the lower Cambrian rocks nowhere exhibit fossils, and hence no living being could have existed in their epoch.

To this there are two replies: the first, that the observational basis of the assertion that the lowest rocks are nowhere fossiliferous is an amazingly small one, seeing how very small an area, in comparison to that of the whole world, has yet been fully searched; the second, that the argument is good for nothing unless the unfossiliferous rocks in question were not only 'contemporaneous' in the geological sense, but 'synchronous' in the chronological sense. To use the 'alibi' illustration again. If a man wishes to prove he was in neither of two places, A and B, on a given day, his witnesses for each place must be prepared to answer for the whole day. If they can only prove that he was not at A in the morning, and not at B in the afternoon, the evidence of his absence from both is 'nil', because he might have been at B in the morning and at A in the afternoon.

Thus everything depends upon the validity of the second assumption. And we must proceed to inquire what is the real meaning of the word "contemporaneous" as employed by geologists. To this end a concrete example may be taken.

The Lias of England and the Lias of Germany, the Cretaceous rocks of Britain and the Cretaceous rocks of Southern India, are termed by geologists "contemporaneous" formations; but whenever any thoughtful geologist is asked whether he means to say that they were deposited synchronously, he says, "No,—only within the same great epoch." And if, in pursuing the inquiry, he is asked what may be the approximate value in time of a "great epoch"—whether it means a hundred years, or a thousand, or a million, or ten million years—his reply is, "I cannot tell."

If the further question be put, whether physical geology is in possession of any method by which the actual synchrony (or the reverse) of any two distant deposits can be ascertained, no such method can be heard of; it being admitted by all the best authorities that neither similarity of mineral composition, nor of physical character, nor even direct continuity of stratum, are 'absolute' proofs of the synchronism of even approximated sedimentary strata: while, for distant deposits, there seems to be no kind of physical evidence attainable of a nature competent to decide whether such deposits were formed simultaneously, or whether they possess any given difference of antiquity. To return to an example already given: All competent authorities will probably assent to the proposition that physical geology does not enable us in any way to reply to this question—Were the British Cretaceous rocks deposited at the same time as those of India, or are they a million of years younger or a million of years older?

Is paleontology able to succeed where physical geology fails? Standard writers on paleontology, as has been seen, assume that she can. They take it for granted, that deposits containing similar organic remains are synchronous—at any rate in a broad sense; and yet, those who will study the eleventh and twelfth chapters of Sir Henry De La Beche's remarkable 'Researches in Theoretical Geology', published now nearly thirty years ago, and will carry out the arguments there most luminously stated, to their logical consequences, may very easily convince themselves that even absolute identity of organic contents is no proof of the synchrony of deposits, while absolute diversity is no proof of difference of date. Sir Henry De La Beche goes even further, and adduces conclusive evidence to show that the different parts of one and the same stratum, having a similar composition throughout, containing the same organic remains, and having similar beds above and below it, may yet differ to any conceivable extent in age.

Edward Forbes was in the habit of asserting that the similarity of the organic contents of distant formations was 'prima facie' evidence, not of their similarity, but of their difference of age; and holding as he did the doctrine of single specific centres, the conclusion was as legitimate as any other; for the two districts must have been occupied by migration from one of the two, or from an intermediate spot, and the chances against exact coincidence of migration and of imbedding are infinite.

In point of fact, however, whether the hypothesis of single or of multiple specific centres be adopted, similarity of organic contents cannot possibly afford any proof of the synchrony of the deposits which contain them; on the contrary, it is demonstrably compatible with the lapse of the most prodigious intervals of time, and with the interposition of vast changes in the organic and inorganic worlds, between the epochs in which such deposits were formed.

On what amount of similarity of their faunae is the doctrine of the contemporaneity of the European and of the North American Silurians based? In the last edition of Sir Charles Lyell's 'Elementary Geology' it is stated, on the authority of a former President of this Society, the late Daniel Sharpe, that between 30 and 40 per cent. of the species of Silurian Mollusca are common to both sides of the Atlantic. By way of due allowance for further discovery, let us double the lesser number and suppose that 60 per cent. of the species are common to the North American and the British Silurians. Sixty per cent. of species in common is, then, proof of contemporaneity.

Now suppose that, a million or two of years hence, when Britain has made another dip beneath the sea and has come up again, some geologist applies this doctrine, in comparing the strata laid bare by the upheaval of the bottom, say, of St. George's Channel with what may then remain of the Suffolk Crag. Reasoning in the same way, he will at once decide the Suffolk Crag and the St. George's Channel beds to be contemporaneous; although we happen to know that a vast period (even in the geological sense) of time, and physical changes of almost unprecedented extent, separate the two.

But if it be a demonstrable fact that strata containing more than 60 or 70 per cent. of species of Mollusca in common, and comparatively close together, may yet be separated by an amount of geological time sufficient to allow of some of the greatest physical changes the world has seen, what becomes of that sort of contemporaneity the sole evidence of which is a similarity of facies, or the identity of half a dozen species, or of a good many genera?

And yet there is no better evidence for the contemporaneity assumed by all who adopt the hypothesis of universal faunae and florae, of a universally uniform climate, and of a sensible cooling of the globe during geological time.

There seems, then, no escape from the admission that neither physical geology, nor paleontology, possesses any method by which the absolute synchronism of two strata can be demonstrated. All that geology can prove is local order of succession. It is mathematically certain that, in any given vertical linear section of an undisturbed series of sedimentary deposits, the bed which lies lowest is the oldest. In many other vertical linear sections of the same series, of course, corresponding beds will occur in a similar order; but, however great may be the probability, no man can say with absolute certainty that the beds in the two sections were synchronously deposited. For areas of moderate extent, it is doubtless true that no practical evil is likely to result from assuming the corresponding beds to be synchronous or strictly contemporaneous; and there are multitudes of accessory circumstances which may fully justify the assumption of such synchrony. But the moment the geologist has to deal with large areas, or with completely separated deposits, the mischief of confounding that "homotaxis" or "similarity of arrangement," which 'can' be demonstrated, with "synchrony" or "identity of date," for which there is not a shadow of proof, under the one common term of "contemporaneity" becomes incalculable, and proves the constant source of gratuitous speculations.

For anything that geology or paleontology are able to show to the contrary, a Devonian fauna and flora in the British Islands may have been contemporaneous with Silurian life in North America, and with a Carboniferous fauna and flora in Africa. Geographical provinces and zones may have been as distinctly marked in the Paleozoic epoch as at present, and those seemingly sudden appearances of new genera and species, which we ascribe to new creation, may be simple results of migration.

It may be so; it may be otherwise. In the present condition of our knowledge and of our methods, one verdict—"not proven, and not provable"—must be recorded against all the grand hypotheses of the paleontologist respecting the general succession of life on the globe. The order and nature of terrestrial life, as a whole, are open questions. Geology at present provides us with most valuable topographical records, but she has not the means of working them into a universal history. Is such a universal history, then, to be regarded as unattainable? Are all the grandest and most interesting problems which offer themselves to the geological student essentially insoluble? Is he in the position of a scientific Tantalus—doomed always to thirst for a knowledge which he cannot obtain? The reverse is to be hoped; nay, it may not be impossible to indicate the source whence help will come.

In commencing these remarks, mention was made of the great obligations under which the naturalist lies to the geologist and paleontologist. Assuredly the time will come when these obligations will be repaid tenfold, and when the maze of the world's past history, through which the pure geologist and the pure paleontologist find no guidance, will be securely threaded by the clue furnished by the naturalist.

All who are competent to express an opinion on the subject are, at present, agreed that the manifold varieties of animal and vegetable form have not either come into existence by chance, nor result from capricious exertions of creative power; but that they have taken place in a definite order, the statement of which order is what men of science term a natural law. Whether such a law is to be regarded as an expression of the mode of operation of natural forces, or whether it is simply a statement of the manner in which a supernatural power has thought fit to act, is a secondary question, so long as the existence of the law and the possibility of its discovery by the human intellect are granted. But he must be a half-hearted philosopher who, believing in that possibility, and having watched the gigantic strides of the biological sciences during the last twenty years, doubts that science will sooner or later make this further step, so as to become possessed of the law of evolution of organic forms—of the unvarying order of that great chain of causes and effects of which all organic forms, ancient and modern, are the links. And then, if ever, we shall be able to begin to discuss, with profit, the questions respecting the commencement of life, and the nature of the successive populations of the globe, which so many seem to think are already answered.

The preceding arguments make no particular claim to novelty; indeed they have been floating more or less distinctly before the minds of geologists for the last thirty years; and if, at the present time, it has seemed desirable to give them more definite and systematic expression, it is because paleontology is every day assuming a greater importance, and now requires to rest on a basis the firmness of which is thoroughly well assured. Among its fundamental conceptions, there must be no confusion between what is certain and what is more or less probable. 2 But, pending the construction of a surer foundation than paleontology now possesses, it may be instructive, assuming for the nonce the general correctness of the ordinary hypothesis of geological contemporaneity, to consider whether the deductions which are ordinarily drawn from the whole body of paleontologic facts are justifiable.

The evidence on which such conclusions are based is of two kinds, negative and positive. The value of negative evidence, in connection with this inquiry, has been so fully and clearly discussed in an address from the chair of this Society 3, which none of us have forgotten, that nothing need at present be said about it; the more, as the considerations which have been laid before you have certainly not tended to increase your estimation of such evidence. It will be preferable to turn to the positive facts of paleontology, and to inquire what they tell us.

We are all accustomed to speak of the number and the extent of the changes in the living population of the globe during geological time as something enormous: and indeed they are so, if we regard only the negative differences which separate the older rocks from the more modern, and if we look upon specific and generic changes as great changes, which from one point of view, they truly are. But leaving the negative differences out of consideration, and looking only at the positive data furnished by the fossil world from a broader point of view—from that of the comparative anatomist who has made the study of the greater modifications of animal form his chief business—a surprise of another kind dawns upon the mind; and under 'this' aspect the smallness of the total change becomes as astonishing as was its greatness under the other.

There are two hundred known orders of plants; of these not one is certainly known to exist exclusively in the fossil state. The whole lapse of geological time has as yet yielded not a single new ordinal type of vegetable structure. 4

The positive change in passing from the recent to the ancient animal world is greater, but still singularly small. No fossil animal is so distinct from those now living as to require to be arranged even in a separate class from those which contain existing forms. It is only when we come to the orders, which may be roughly estimated at about a hundred and thirty, that we meet with fossil animals so distinct from those now living as to require orders for themselves; and these do not amount, on the most liberal estimate, to more than about 10 per cent. of the whole.

There is no certainly known extinct order of Protozoa; there is but one among the Coelenterata—that of the rugose corals; there is none among the Mollusca; there are three, the Cystidea, Blastoidea, and Edrioasterida, among the Echinoderms; and two, the Trilobita and Eurypterida, among the Crustacea; making altogether five for the great sub-kingdom of Annulosa. Among Vertebrates there is no ordinally distinct fossil fish: there is only one extinct order of Amphibia—the Labyrinthodonts; but there are at least four distinct orders of Reptilia, viz. the Ichthyosauria, Plesiosauria, Pterosauria, Dinosauria, and perhaps another or two. There is no known extinct order of Birds, and no certainly known extinct order of Mammals, the ordinal distinctness of the "Toxodontia" being doubtful.

The objection that broad statements of this kind, after all, rest largely on negative evidence is obvious, but it has less force than may at first be supposed; for, as might be expected from the circumstances of the case, we possess more abundant positive evidence regarding Fishes and marine Mollusks than respecting any other forms of animal life; and yet these offer us, through the whole range of geological time, no species ordinally distinct from those now living; while the far less numerous class of Echinoderms presents three; and the Crustacea two, such orders, though none of these come down later than the Paleozoic age. Lastly, the Reptilia present the extraordinary and exceptional phenomenon of as many extinct as existing orders, if not more; the four mentioned maintaining their existence from the Lias to the Chalk inclusive.

Some years ago one of your Secretaries pointed out another kind of positive paleontologic evidence tending towards the same conclusion—afforded by the existence of what he termed "persistent types" of vegetable and of animal life. 5 He stated, on the authority of Dr. Hooker, that there are Carboniferous plants which appear to be generically identical with some now living; that the cone of the Oolitic 'Araucaria' is hardly distinguishable from that of an existing species; that a true 'Pinus' appears in the Purbecks, and a 'Juglans' in the Chalk; while, from the Bagshot Sands, a 'Banksia', the wood of which is not distinguishable from that of species now living in Australia, had been obtained.

Turning to the animal kingdom, he affirmed the tabulate corals of the Silurian rocks to be wonderfully like those which now exist; while even the families of the Aporosa were all represented in the older Mesozoic rocks.

Among the Molluska similar facts were adduced. Let it be borne in mind that 'Avicula', 'Mytails', 'Chiton', 'Natica', 'Patella', 'Trochus', 'Discina', 'Orbicula', 'Lingula', 'Rhynchonella', and 'Nautilus', all of which are existing 'genera', are given without a doubt as Silurian in the last edition of 'Siluria'; while the highest forms of the highest Cephalopods are represented in the Lias by a genus, 'Belemnoteuthis', which presents the closest relation to the existing 'Loligo'.

The two highest groups of the Annulosa, the Insecta and the Arachnida, are represented in the Coal, either by existing genera, or by forms differing from existing genera in quite minor peculiarities.

Turning to the Vertebrata, the only Paleozoic Elasmobranch Fish of which we have any complete knowledge is the Devonian and Carboniferous 'Pleuracanthus', which differs no more from existing Sharks than these do from one another.

Again, vast as is the number of undoubtedly Ganoid fossil Fishes, and great as is their range in time, a large mass of evidence has recently been adduced to show that almost all those respecting which we possess sufficient information, are referable to the same sub-ordinal groups as the existing 'Lepidosteus', 'Polypterus', and Sturgeon; and that a singular relation obtains between the older and the younger Fishes; the former, the Devonian Ganoids, being almost all members of the same sub-order as 'Polypterus', while the Mesozoic Ganoids are almost all similarly allied to 'Lepidosteus'. 6

Again, what can be more remarkable than the singular constancy of structure preserved throughout a vast period of time by the family of the Pycnodonts and by that of the true Coelacanths; the former persisting, with but insignificant modifications, from the Carboniferous to the Tertiary rocks, inclusive; the latter existing, with still less change, from the Carboniferous rocks to the Chalk, inclusive?

Among Reptiles, the highest living group, that of the Crocodilia, is represented, at the early part of the Mesozoic epoch, by species identical in the essential characters of their organization with those now living, and differing from the latter only in such matters as the form of the articular facets of the vertebral centra, in the extent to which the nasal passages are separated from the cavity of the mouth by bone, and in the proportions of the limbs.

And even as regards the Mammalia, the scanty remains of Triassic and Oolitic species afford no foundation for the supposition that the organization of the oldest forms differed nearly so much from some of those which now live as these differ from one another.

It is needless to multiply these instances; enough has been said to justify the statement that, in view of the immense diversity of known animal and vegetable forms, and the enormous lapse of time indicated by the accumulation of fossiliferous strata, the only circumstance to be wondered at is, not that the changes of life, as exhibited by positive evidence, have been so great, but that they have been so small.

Be they great or small, however, it is desirable to attempt to estimate them. Let us, therefore, take each great division of the animal world in succession, and, whenever an order or a family can be shown to have had a prolonged existence, let us endeavour to ascertain how far the later members of the group differ from the earlier ones. If these later members, in all or in many cases, exhibit a certain amount of modification, the fact is, so far, evidence in favour of a general law of change; and, in a rough way, the rapidity of that change will be measured by the demonstrable amount of modification. On the other hand, it must be recollected that the absence of any modification, while it may leave the doctrine of the existence of a law of change without positive support, cannot possibly disprove all forms of that doctrine, though it may afford a sufficient refutation of any of them.

The PROTOZOA.—The Protozoa are represented throughout the whole range of geological series, from the Lower Silurian formation to the present day. The most ancient forms recently made known by Ehrenberg are exceedingly like those which now exist: no one has ever pretended that the difference between any ancient and any modern Foraminifera is of more than generic value, nor are the oldest Foraminifera either simpler, more embryonic, or less differentiated, than the existing forms.

The COELENTERATA.—The Tabulate Corals have existed from the Silurian epoch to the present day, but I am not aware that the ancient 'Heliolites' possesses a single mark of a more embryonic or less differentiated character, or less high organization, than the existing 'Heliopora'. As for the Aporose Corals, in what respect is the Silurian 'Paleocyclus' less highly organized or more embryonic than the modern 'Fungia', or the Liassic Aporosa than the existing members of the same families?

The 'Mollusca'.—In what sense is the living 'Waldheimia' less embryonic, or more specialized; than the paleozoic 'Spirifer'; or the existing 'Rhynchonellae', 'Craniae', 'Discinae', 'Lingulae', than the Silurian species of the same genera? In what sense can 'Loligo' or 'Spirula' be said to be more specialized, or less embryonic, than 'Belemnites'; or the modern species of Lamellibranch and Gasteropod genera, than the Silurian species of the same genera?

The ANNULOSA.—The Carboniferous Insecta and Arachnida are neither less specialized, nor more embryonic, than these that now live, nor are the Liassic Cirripedia and Macrura; while several of the Brachyura, which appear in the Chalk, belong to existing genera; and none exhibit either an intermediate, or an embryonic, character.

The VERTEBRARA.—Among fishes I have referred to the Coelacanthini (comprising the genera 'Coelacanthus', 'Holophagus', 'Undina', and 'Macropoma') as affording an example of a persistent type; and it is most remarkable to note the smallness of the differences between any of these fishes (affecting at most the proportions of the body and fins, and the character and sculpture of the scales), notwithstanding their enormous range in time. In all the essentials of its very peculiar structure, the 'Macropoma' of the Chalk is identical with the 'Coelacanthus' of the Coal. Look at the genus 'Lepidotus', again, persisting without a modification of importance from the Liassic to the Eocene formations inclusive.

Or among the Teleostei—in what respect is the 'Beryx' of the Chalk more embryonic, or less differentiated, than 'Beryx lineatus' of King George's Sound?

Or to turn to the higher Vertebrata—in what sense are the Liassic Chelonia inferior to those which now exist? How are the Cretaceous Ichthyosauria, Plesiosauria, or Pterosauria less embryonic, or more differentiated, species than those of the Lias?

Or lastly, in what circumstance is the 'Phascolotherium' more embryonic, or of a more generalized type, than the modern Opossum; or a 'Lophiodon', or a 'Paleotherium', than a modern 'Tapirus' or 'Hyrax'?

These examples might be almost indefinitely multiplied, but surely they are sufficient to prove that the only safe and unquestionable testimony we can procure—positive evidence—fails to demonstrate any sort of progressive modification towards a less embryonic, or less generalised, type in a great many groups of animals of long-continued geological existence. In these groups there is abundant evidence of variation—none of what is ordinarily understood as progression; and, if the known geological record is to be regarded as even any considerable fragment of the whole, it is inconceivable that any theory of a necessarily progressive development can stand, for the numerous orders and families cited afford no trace of such a process.

But it is a most remarkable fact, that, while the groups which have been mentioned, and many besides, exhibit no sign of progressive modification, there are others, co-existing with them, under the same conditions, in which more or less distinct indications of such a process seems to be traceable. Among such indications I may remind you of the predominance of Holostome Gasteropoda in the older rocks as compared with that of Siphonostome Gasteropoda in the later. A case less open to the objection of negative evidence, however, is that afforded by the Tetrabranchiate Cephalopoda, the forms of the shells and of the septal sutures exhibiting a certain increase of complexity in the newer genera. Here, however, one is met at once with the occurrence of 'Orthoceras' and 'Baculites' at the two ends of the series, and of the fact that one of the simplest Genera, 'Nautilus', is that which now exists.

The Crinoidea, in the abundance of stalked forms in the ancient formations as compared with their present rarity, seem to present us with a fair case of modification from a more embryonic towards a less embryonic condition. But then, on careful consideration of the facts, the objection arises that the stalk, calyx, and arms of the paleozoic Crinoid are exceedingly different from the corresponding organs of a larval 'Comatula'; and it might with perfect justice be argued that 'Actinocrinus' and 'Eucalyptocrinus', for example, depart to the full as widely, in one direction, from the stalked embryo of 'Comatula', as 'Comatula' itself does in the other.

The Echinidea, again, are frequently quoted as exhibiting a gradual passage from a more generalized to a more specialized type, seeing that the elongated, or oval, Spatangoids appear after the spheroidal Echinoids. But here it might be argued, on the other hand, that the spheroidal Echinoids, in reality, depart further from the general plan and from the embryonic form than the elongated Spatangoids do; and that the peculiar dental apparatus and the pedicellariae of the former are marks of at least as great differentiation as the petaloid ambulacra and semitae of the latter.

Once more, the prevalence of Macrurous before Brachyurous Podophthalmia is, apparently, a fair piece of evidence in favour of progressive modification in the same order of Crustacea; and yet the case will not stand much sifting, seeing that the Macrurous Podophthalmia depart as far in one direction from the common type of Podophthalmia, or from any embryonic condition of the Brachyura, as the Brachyura do in the other; and that the middle terms between Macrura and Brachyura—the Anomura—are little better represented in the older Mesozoic rocks than the Brachyura are.

None of the cases of progressive modification which are cited from among the Invertebrata appear to me to have a foundation less open to criticism than these; and if this be so, no careful reasoner would, I think, be inclined to lay very great stress upon them. Among the Vertebrata, however, there are a few examples which appear to be far less open to objection.

It is, in fact, true of several groups of Vertebrata which have lived through a considerable range of time, that the endoskeleton (more particularly the spinal column) of the older genera presents a less ossified, and, so far, less differentiated, condition than that of the younger genera. Thus the Devonian Ganoids, though almost all members of the same sub-order as 'Polypterus', and presenting numerous important resemblances to the existing genus, which possesses biconcave vertebrae, are, for the most part, wholly devoid of ossified vertebral centra. The Mesozoic Lepidosteidae, again, have, at most, biconcave vertebrae, while the existing 'Lepidosteus' has Salamandroid, opisthocoelous, vertebrae. So, none of the Paleozoic Sharks have shown themselves to be possessed of ossified vertebrae, while the majority of modern Sharks possess such vertebrae. Again, the more ancient Crocodilia and Lacertilia have vertebrae with the articular facets of their centra flattened or biconcave, while the modern members of the same group have them procoelous. But the most remarkable examples of progressive modification of the vertebral column, in correspondence with geological age, are those afforded by the Pycnodonts among fish, and the Labyrinthodonts among Amphibia.

The late able ichthyologist Heckel pointed out the fact, that, while the Pycnodonts never possess true vertebral centra, they differ in the degree of expansion and extension of the ends of the bony arches of the vertebrae upon the sheath of the notochord; the Carboniferous forms exhibiting hardly any such expansion, while the Mesozoic genera present a greater and greater development, until, in the Tertiary forms, the expanded ends become suturally united so as to form a sort of false vertebra. Hermann von Meyer, again, to whose luminous researches we are indebted for our present large knowledge of the organization of the older Labyrinthodonts, has proved that the Carboniferous 'Archegosaurus' had very imperfectly developed vertebral centra, while the Triassic 'Mastodonsaurus' had the same parts completely ossified. 7

The regularity and evenness of the dentition of the 'Anoplotherium', as contrasted with that of existing Artiodactyles, and the assumed nearer approach of the dentition of certain ancient Carnivores to the typical arrangement, have also been cited as exemplifications of a law of progressive development, but I know of no other cases based on positive evidence which are worthy of particular notice.

What, then, does an impartial survey of the positively ascertained truths of paleontology testify in relation to the common doctrines of progressive modification, which suppose that modification to have taken place by a necessary progress from more to less embryonic forms, or from more to less generalized types, within the limits of the period represented by the fossiliferous rocks?

It negatives those doctrines; for it either shows us no evidence of any such modification, or demonstrates it to have been very slight; and as to the nature of that modification, it yields no evidence whatsoever that the earlier members of any long-continued group were more generalized in structure than the later ones. To a certain extent, indeed, it may be said that imperfect ossification of the vertebral column is an embryonic character; but, on the other hand, it would be extremely incorrect to suppose that the vertebral columns of the older Vertebrata are in any sense embryonic in their whole structure.

Obviously, if the earliest fossiliferous rocks now known are coeval with the commencement of life, and if their contents give us any just conception of the nature and the extent of the earliest fauna and flora, the insignificant amount of modification which can be demonstrated to have taken place in any one group of animals, or plants, is quite incompatible with the hypothesis that all living forms are the results of a necessary process of progressive development, entirely comprised within the time represented by the fossiliferous rocks.

Contrariwise, any admissible hypothesis of progressive modification must be compatible with persistence without progression, through indefinite periods. And should such an hypothesis eventually be proved to be true, in the only way in which it can be demonstrated, viz. by observation and experiment upon the existing forms of life, the conclusion will inevitably present itself, that the Paleozoic, Mesozoic, and Cainozoic faunae and florae, taken together, bear somewhat the same proportion to the whole series of living beings which have occupied this globe, as the existing fauna and flora do to them.

Such are the results of paleontology as they appear, and have for some years appeared, to the mind of an inquirer who regards that study simply as one of the applications of the great biological sciences, and who desires to see it placed upon the same sound basis as other branches of physical inquiry. If the arguments which have been brought forward are valid, probably no one, in view of the present state of opinion, will be inclined to think the time wasted which has been spent upon their elaboration.

1 (return)
[ The Anniversary Address to the Geological Society for 1862.]

2 (return)
[ "le plus grand service qu'on puisse rendre a la science est d'y faire place nette avant d'y rien construire."—CUVIER]

3 (return)
[ Anniversary Address for 1851, 'Quart. Journ. Geol. Soc.' vol. vii.]

4 (return)
[ See Hooker's 'Introductory Essay to the Flora of Tasmania', p. xxiii.]

5 (return)
[ See the abstract of a Lecture "On the Persistent Types of Animal Life," in the 'Notices of the Meetings of the Royal Institution of Great Britain'.—June 3, 1859, vol. iii. p. 151.

6 (return)
[ "Memoirs of the Geological Survey of the United Kingdom.—Decade x. Preliminary Essay upon the Systematic Arrangement of the Fishes of the Devonian Epoch."]

7 (return)
[ As the Address is passing through the press (March 7, 1862), evidence lies before me of the existence of a new Labyrinthodont ('Pholidogaster'), from the Edinburgh coal-field, with well-ossified vertebral centra.]

*** END OF THE PROJECT GUTENBERG EBOOK GEOLOGICAL CONTEMPORANEITY AND PERSISTENT TYPES OF LIFE ***

Updated editions will replace the previous one—the old editions will be renamed.

Creating the works from print editions not protected by U.S. copyright law means that no one owns a United States copyright in these works, so the Foundation (and you!) can copy and distribute it in the United States without permission and without paying copyright royalties. Special rules, set forth in the General Terms of Use part of this license, apply to copying and distributing Project Gutenberg™ electronic works to protect the PROJECT GUTENBERG™ concept and trademark. Project Gutenberg is a registered trademark, and may not be used if you charge for an eBook, except by following the terms of the trademark license, including paying royalties for use of the Project Gutenberg trademark. If you do not charge anything for copies of this eBook, complying with the trademark license is very easy. You may use this eBook for nearly any purpose such as creation of derivative works, reports, performances and research. Project Gutenberg eBooks may be modified and printed and given away—you may do practically ANYTHING in the United States with eBooks not protected by U.S. copyright law. Redistribution is subject to the trademark license, especially commercial redistribution.

START: FULL LICENSE

THE FULL PROJECT GUTENBERG™ LICENSE

PLEASE READ THIS BEFORE YOU DISTRIBUTE OR USE THIS WORK

To protect the Project Gutenberg™ mission of promoting the free distribution of electronic works, by using or distributing this work (or any other work associated in any way with the phrase “Project Gutenberg”), you agree to comply with all the terms of the Full Project Gutenberg License available with this file or online at www.gutenberg.org/license.

Section 1. General Terms of Use and Redistributing Project Gutenberg electronic works

1.A. By reading or using any part of this Project Gutenberg electronic work, you indicate that you have read, understand, agree to and accept all the terms of this license and intellectual property (trademark/copyright) agreement. If you do not agree to abide by all the terms of this agreement, you must cease using and return or destroy all copies of Project Gutenberg electronic works in your possession. If you paid a fee for obtaining a copy of or access to a Project Gutenberg electronic work and you do not agree to be bound by the terms of this agreement, you may obtain a refund from the person or entity to whom you paid the fee as set forth in paragraph 1.E.8.

1.B. “Project Gutenberg” is a registered trademark. It may only be used on or associated in any way with an electronic work by people who agree to be bound by the terms of this agreement. There are a few things that you can do with most Project Gutenberg electronic works even without complying with the full terms of this agreement. See paragraph 1.C below. There are a lot of things you can do with Project Gutenberg electronic works if you follow the terms of this agreement and help preserve free future access to Project Gutenberg electronic works. See paragraph 1.E below.

1.C. The Project Gutenberg Literary Archive Foundation (“the Foundation” or PGLAF), owns a compilation copyright in the collection of Project Gutenberg electronic works. Nearly all the individual works in the collection are in the public domain in the United States. If an individual work is unprotected by copyright law in the United States and you are located in the United States, we do not claim a right to prevent you from copying, distributing, performing, displaying or creating derivative works based on the work as long as all references to Project Gutenberg are removed. Of course, we hope that you will support the Project Gutenberg mission of promoting free access to electronic works by freely sharing Project Gutenberg works in compliance with the terms of this agreement for keeping the Project Gutenberg name associated with the work. You can easily comply with the terms of this agreement by keeping this work in the same format with its attached full Project Gutenberg License when you share it without charge with others.

1.D. The copyright laws of the place where you are located also govern what you can do with this work. Copyright laws in most countries are in a constant state of change. If you are outside the United States, check the laws of your country in addition to the terms of this agreement before downloading, copying, displaying, performing, distributing or creating derivative works based on this work or any other Project Gutenberg work. The Foundation makes no representations concerning the copyright status of any work in any country other than the United States.

1.E. Unless you have removed all references to Project Gutenberg:

1.E.1. The following sentence, with active links to, or other immediate access to, the full Project Gutenberg License must appear prominently whenever any copy of a Project Gutenberg work (any work on which the phrase “Project Gutenberg” appears, or with which the phrase “Project Gutenberg” is associated) is accessed, displayed, performed, viewed, copied or distributed:

This eBook is for the use of anyone anywhere in the United States and most other parts of the world at no cost and with almost no restrictions whatsoever. You may copy it, give it away or re-use it under the terms of the Project Gutenberg™ License included with this eBook or online at www.gutenberg.org. If you are not located in the United States, you will have to check the laws of the country where you are located before using this eBook.

1.E.2. If an individual Project Gutenberg electronic work is derived from texts not protected by U.S. copyright law (does not contain a notice indicating that it is posted with permission of the copyright holder), the work can be copied and distributed to anyone in the United States without paying any fees or charges. If you are redistributing or providing access to a work with the phrase “Project Gutenberg” associated with or appearing on the work, you must comply either with the requirements of paragraphs 1.E.1 through 1.E.7 or obtain permission for the use of the work and the Project Gutenberg trademark as set forth in paragraphs 1.E.8 or 1.E.9.

1.E.3. If an individual Project Gutenberg electronic work is posted with the permission of the copyright holder, your use and distribution must comply with both paragraphs 1.E.1 through 1.E.7 and any additional terms imposed by the copyright holder. Additional terms will be linked to the Project Gutenberg License for all works posted with the permission of the copyright holder found at the beginning of this work.

1.E.4. Do not unlink or detach or remove the full Project Gutenberg License terms from this work, or any files containing a part of this work or any other work associated with Project Gutenberg.

1.E.5. Do not copy, display, perform, distribute or redistribute this electronic work, or any part of this electronic work, without prominently displaying the sentence set forth in paragraph 1.E.1 with active links or immediate access to the full terms of the Project Gutenberg License.

1.E.6. You may convert to and distribute this work in any binary, compressed, marked up, nonproprietary or proprietary form, including any word processing or hypertext form. However, if you provide access to or distribute copies of a Project Gutenberg work in a format other than “Plain Vanilla ASCII” or other format used in the official version posted on the official Project Gutenberg website (www.gutenberg.org), you must, at no additional cost, fee or expense to the user, provide a copy, a means of exporting a copy, or a means of obtaining a copy upon request, of the work in its original “Plain Vanilla ASCII” or other form. Any alternate format must include the full Project Gutenberg License as specified in paragraph 1.E.1.

1.E.7. Do not charge a fee for access to, viewing, displaying, performing, copying or distributing any Project Gutenberg works unless you comply with paragraph 1.E.8 or 1.E.9.

1.E.8. You may charge a reasonable fee for copies of or providing access to or distributing Project Gutenberg electronic works provided that:

- • You pay a royalty fee of 20% of the gross profits you derive from the use of Project Gutenberg works calculated using the method you already use to calculate your applicable taxes. The fee is owed to the owner of the Project Gutenberg trademark, but he has agreed to donate royalties under this paragraph to the Project Gutenberg Literary Archive Foundation. Royalty payments must be paid within 60 days following each date on which you prepare (or are legally required to prepare) your periodic tax returns. Royalty payments should be clearly marked as such and sent to the Project Gutenberg Literary Archive Foundation at the address specified in Section 4, “Information about donations to the Project Gutenberg Literary Archive Foundation.”

- • You provide a full refund of any money paid by a user who notifies you in writing (or by e-mail) within 30 days of receipt that s/he does not agree to the terms of the full Project Gutenberg™ License. You must require such a user to return or destroy all copies of the works possessed in a physical medium and discontinue all use of and all access to other copies of Project Gutenberg™ works.

- • You provide, in accordance with paragraph 1.F.3, a full refund of any money paid for a work or a replacement copy, if a defect in the electronic work is discovered and reported to you within 90 days of receipt of the work.

- • You comply with all other terms of this agreement for free distribution of Project Gutenberg™ works.

1.E.9. If you wish to charge a fee or distribute a Project Gutenberg™ electronic work or group of works on different terms than are set forth in this agreement, you must obtain permission in writing from the Project Gutenberg Literary Archive Foundation, the manager of the Project Gutenberg™ trademark. Contact the Foundation as set forth in Section 3 below.

1.F.

1.F.1. Project Gutenberg volunteers and employees expend considerable effort to identify, do copyright research on, transcribe and proofread works not protected by U.S. copyright law in creating the Project Gutenberg™ collection. Despite these efforts, Project Gutenberg™ electronic works, and the medium on which they may be stored, may contain “Defects,” such as, but not limited to, incomplete, inaccurate or corrupt data, transcription errors, a copyright or other intellectual property infringement, a defective or damaged disk or other medium, a computer virus, or computer codes that damage or cannot be read by your equipment.

1.F.2. LIMITED WARRANTY, DISCLAIMER OF DAMAGES - Except for the “Right of Replacement or Refund” described in paragraph 1.F.3, the Project Gutenberg Literary Archive Foundation, the owner of the Project Gutenberg™ trademark, and any other party distributing a Project Gutenberg™ electronic work under this agreement, disclaim all liability to you for damages, costs and expenses, including legal fees. YOU AGREE THAT YOU HAVE NO REMEDIES FOR NEGLIGENCE, STRICT LIABILITY, BREACH OF WARRANTY OR BREACH OF CONTRACT EXCEPT THOSE PROVIDED IN PARAGRAPH 1.F.3. YOU AGREE THAT THE FOUNDATION, THE TRADEMARK OWNER, AND ANY DISTRIBUTOR UNDER THIS AGREEMENT WILL NOT BE LIABLE TO YOU FOR ACTUAL, DIRECT, INDIRECT, CONSEQUENTIAL, PUNITIVE OR INCIDENTAL DAMAGES EVEN IF YOU GIVE NOTICE OF THE POSSIBILITY OF SUCH DAMAGE.

1.F.3. LIMITED RIGHT OF REPLACEMENT OR REFUND - If you discover a defect in this electronic work within 90 days of receiving it, you can receive a refund of the money (if any) you paid for it by sending a written explanation to the person you received the work from. If you received the work on a physical medium, you must return the medium with your written explanation. The person or entity that provided you with the defective work may elect to provide a replacement copy in lieu of a refund. If you received the work electronically, the person or entity providing it to you may choose to give you a second opportunity to receive the work electronically in lieu of a refund. If the second copy is also defective, you may demand a refund in writing without further opportunities to fix the problem.

1.F.4. Except for the limited right of replacement or refund set forth in paragraph 1.F.3, this work is provided to you ‘AS-IS’, WITH NO OTHER WARRANTIES OF ANY KIND, EXPRESS OR IMPLIED, INCLUDING BUT NOT LIMITED TO WARRANTIES OF MERCHANTABILITY OR FITNESS FOR ANY PURPOSE.

1.F.5. Some states do not allow disclaimers of certain implied warranties or the exclusion or limitation of certain types of damages. If any disclaimer or limitation set forth in this agreement violates the law of the state applicable to this agreement, the agreement shall be interpreted to make the maximum disclaimer or limitation permitted by the applicable state law. The invalidity or unenforceability of any provision of this agreement shall not void the remaining provisions.

1.F.6. INDEMNITY - You agree to indemnify and hold the Foundation, the trademark owner, any agent or employee of the Foundation, anyone providing copies of Project Gutenberg™ electronic works in accordance with this agreement, and any volunteers associated with the production, promotion and distribution of Project Gutenberg™ electronic works, harmless from all liability, costs and expenses, including legal fees, that arise directly or indirectly from any of the following which you do or cause to occur: (a) distribution of this or any Project Gutenberg work, (b) alteration, modification, or additions or deletions to any Project Gutenberg work, and (c) any Defect you cause.

Section 2. Information about the Mission of Project Gutenberg

Project Gutenberg is synonymous with the free distribution of electronic works in formats readable by the widest variety of computers including obsolete, old, middle-aged and new computers. It exists because of the efforts of hundreds of volunteers and donations from people in all walks of life.

Volunteers and financial support to provide volunteers with the assistance they need are critical to reaching Project Gutenberg’s goals and ensuring that the Project Gutenberg collection will remain freely available for generations to come. In 2001, the Project Gutenberg Literary Archive Foundation was created to provide a secure and permanent future for Project Gutenberg and future generations. To learn more about the Project Gutenberg Literary Archive Foundation and how your efforts and donations can help, see Sections 3 and 4 and the Foundation information page at www.gutenberg.org.

Section 3. Information about the Project Gutenberg Literary Archive Foundation

The Project Gutenberg Literary Archive Foundation is a non-profit 501(c)(3) educational corporation organized under the laws of the state of Mississippi and granted tax exempt status by the Internal Revenue Service. The Foundation’s EIN or federal tax identification number is 64-6221541. Contributions to the Project Gutenberg Literary Archive Foundation are tax deductible to the full extent permitted by U.S. federal laws and your state’s laws.

The Foundation’s business office is located at 41 Watchung Plaza #516, Montclair NJ 07042, USA, +1 (862) 621-9288. Email contact links and up to date contact information can be found at the Foundation’s website and official page at www.gutenberg.org/contact

Section 4. Information about Donations to the Project Gutenberg Literary Archive Foundation

Project Gutenberg™ depends upon and cannot survive without widespread public support and donations to carry out its mission of increasing the number of public domain and licensed works that can be freely distributed in machine-readable form accessible by the widest array of equipment including outdated equipment. Many small donations ($1 to $5,000) are particularly important to maintaining tax exempt status with the IRS.

The Foundation is committed to complying with the laws regulating charities and charitable donations in all 50 states of the United States. Compliance requirements are not uniform and it takes a considerable effort, much paperwork and many fees to meet and keep up with these requirements. We do not solicit donations in locations where we have not received written confirmation of compliance. To SEND DONATIONS or determine the status of compliance for any particular state visit www.gutenberg.org/donate.

While we cannot and do not solicit contributions from states where we have not met the solicitation requirements, we know of no prohibition against accepting unsolicited donations from donors in such states who approach us with offers to donate.

International donations are gratefully accepted, but we cannot make any statements concerning tax treatment of donations received from outside the United States. U.S. laws alone swamp our small staff.

Please check the Project Gutenberg web pages for current donation methods and addresses. Donations are accepted in a number of other ways including checks, online payments and credit card donations. To donate, please visit: www.gutenberg.org/donate.

Section 5. General Information About Project Gutenberg electronic works

Professor Michael S. Hart was the originator of the Project Gutenberg concept of a library of electronic works that could be freely shared with anyone. For forty years, he produced and distributed Project Gutenberg eBooks with only a loose network of volunteer support.

Project Gutenberg eBooks are often created from several printed editions, all of which are confirmed as not protected by copyright in the U.S. unless a copyright notice is included. Thus, we do not necessarily keep eBooks in compliance with any particular paper edition.

Most people start at our website which has the main PG search facility: www.gutenberg.org.

This website includes information about Project Gutenberg, including how to make donations to the Project Gutenberg Literary Archive Foundation, how to help produce our new eBooks, and how to subscribe to our email newsletter to hear about new eBooks.

Table of Contents

- GEOLOGICAL CONTEMPORANEITY AND PERSISTENT TYPES OF LIFE.

- THE FULL PROJECT GUTENBERG™ LICENSE